On cars using speed density or volumetric efficiency engine mangement, the
IAT needs to basically be measuring the air temp as it enters the engine. (where the MAP sensor is. As oposed to where the
MAF sensor is on
MAF based engine management)
Speed density tuning and VE tuning are similar, in that the
ECU calculates the air mass entering the engine by knowing its pressure and its temperature. (And knowing the engines displacement - volume) It basically uses the universal laws of gas to work it out. (Temperature / pressure / volume / mass. If you know 3, you can always work out the 4th.)
Link G4+ ECUs call VE tuning "modelled" and standard speed density as "traditional".
Im not sure if Jakes is set up using traditional or modelled. But they kinda operate the same. Modelled you basically feed it all the info including target AFR, and only tune a "VE table", and the
ECU calculates the rest. Tradittional has a fueling fuel table which is what will be tuned.